Chapter 31: Plant Reproduction

31.1 Reproductive Strctures of Flowering Plants

31.1.1 Think Sporophyte and Gametophyte


Know the overview of the flowering plant life cycle. Fig. 31.3

The Sporophyte (diploid) stage is the dominant stage producing microscopic spores that produce the gametophytes. The male gametophyte (haploid) is the pollen grain that is dispersed but the female gametophyte (haploid) lives parasitically in the sporophyte flower tissue.

31.1.2 Components of Flowers


Know the components of a flower. Fig. 31.3

The flower in Fig. 31.3 is a complete flower with, male and female components and other structures given. Flowers may only have components of this complete design. For example: Some plant species will have male flowers and one plant and female flowers on another plant.

31.1.3 Where Pollen ad Eggs Develop


The male gametophyte, the pollen grains develop in the athers, a part of the stamen structure.

The female gametophyte develops in the ovary of the carpel.

Flowers with both male and female structures are called perfect flowers and others are imperfect flowers containing male or female structures.

 

31.2 Pollen Sets Me Sneezing

Note: Be able to recall (in human cells) the cell production of mucus in response to the presence of pollen on the surface of the cell.

Take a look at the surface of the pollen in Fig. 31.5.

Different pollen surfaces are specific to the plant species that produced it.

31.3 A New Generation Begins

Be able to summarize the complete process in Fig. 31.6 including double fertilization.

31.3.1 From Microspores to Pollen Grains

Cells in the pollen sacs of the anther divide by meiosis and produce haploid microspores. These become the pollen grains. Sperm nuclei form in the pollen grain that will travel through the pollen tube to fertilize tissue in the female gametophyte.

31.3.2 From Megaspores to Eggs

In the ovules of the ovary, meoitic cell divisions form haploid megaspores. Further mitotic divisions of this haploid tissue leads to the embryo sac, the female gametophyte. Within the embryo sac, one cell is the egg and another cell with two nuclei, is the cell that will develop into the endsperm, the nutritive tissue for the embryo as it develops.

31.3.3 From Pollination to Fertilization


Pollination is when the pollen grains are transfered to the stigma.

Fertilization occurs when sperm nuclei fuse with cells of the embryo sac. Double fertilization occurs:

1. pollen tube and nucleus with two sperm cells penetrate the embryo sac.

2. the sperm nuclei fertilize both: (therefore-double fertilization)

a. egg is fertilized by the sperm nuclei, the resulting zygote is a diploid embryo.

b. two nuclei endosperm mother cell will produce the endosperm nutritive tissue.

Know Fig. 31.6 and be sure to read: A Coevolutionary Tale - Commentary of Theory

 

31.4 From Zygote to Seeds and Fruits

31.4.1 Formation of the Embryo Sporophyte Fig. 31.7

The embryo of a dicot develops codyledons, or seed leaves. The endosperm nutritive materials are transfered into these codyledons for storage until germination.

In moncots the endosperm is left until germination, when the codyledon transfers enzymes to the endosperm to break down and distribute the nutrients of the endosperm to the growing seedling.

Be able to locate the embryo and the cotyledons - is it a monocot or dicot??

31.4.2 Formation of Seeds and Fruits


Seed is the mature ovule containing the embryo, the food reserves, and the seed coat.

The seed is a protected area for embryo development.

The ovule's integument thickens and hardens into a seed coat.

The fruit develops from the ovary and assoicated tissue around the seeds.

Fruit descriptions involve fruits that dry or fleshy, patterns of the incorporation of ovaries and associated tissue (note: one ovary - simple, or multiple ovaries - compound), and flower combinations (pineapples).Fig. 31.8

Examples: Table 31.1 - Fleshy (apple) or dry & hard (walnut)

Fleshy fruit has three divisions: endocarp (tissue around the seed[s]), mesocarp (middle, fleshy part), and the exocarp (skin) [All together refered to as: pericarp]

31.5 Dispersal of Fruits and Seeds

Modifications specific for each plant are linked to different means of seed dispersion. Fruit function as the means of seed dispersion.

Types of dispersal:
1. wind-dispersed fruit - (The pericarp might be extended like "wings" to catch the wind.)
2. water-dispersed fruit (example: coconuts)
3. animal-dispersed fruit - eaten by animals and the seeds coats are only slightly digested and thus dispersed by the animal in its feces and prepared for germination.

31.6 Why So Many Flowers and So Few Fruits?

You should be able to put forth an hypothesis for this question after reading this section presenting the flowering of the giant saguaro in Arizona.

31.7 Asexual Reproduction of Flowering Plants

31.7.1 Asexual Reproduction in Nature

Plants can reproduce without fertilization through various modes of vegetative growth. Table 31.2 This is the result of mitotic divisions of plant parts (on the plant or separated from the plant) leading to the production of a complete individual, a clone of the parent plant. Fig. 31.11

Parthenogenesis - the embryo develops from an unfertilized egg. It is prompted by the presence of pollen on the stigma, and chemicals may prompt the fusing of two eggs in the embryo sac to initiate the embryo development.

31.7.2 Induced Propagation

Tissue taken from a plant and treated with growth factors, can be induced to produce a mass of dividing cells that can result in vegetative reproduction. Even differentiated cells can be induced. Induction often involves exposing the plant tissue to growth promoting chemicals that are in the plant naturally but are here exposed to tissues that are prompted to produce a new plant. Fig. 31.12